Department of Plant Nutrition, Campus Universitario de Espinardo, CEBAS-CSIC, Ed 25, Espinardo, Murcia, 30100, Spain. Division of Plant Sciences and Interdisciplinary Plant Group, College of Agriculture, Food and Natural Resources, Christopher S. Bond Life Sciences Center, University of Missouri, 1201 Rollins Street, Columbia, MO, 65201, USA. Department of Plant Sciences, University of California, 1 Shields Avenue, Davis, CA, 95616, USA. Departamento de Ciencias Agrarias y del Medio Natural, Universitat Jaume I, Av. de Vicent Sos Baynat, s/n, Castelló de la Plana, 12071, Spain.